A Quote by Charles Darwin
I have called this principle, by which each slight variation, if useful, is preserved, by the term of Natural Selection.
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I have called this principle, by which each slight variation, if useful, is preserved, by the term Natural Selection, in order to mark its relation to man's power of selection. But the expression often used by Mr. Herbert Spencer of the Survival of the Fittest is more accurate, and is sometimes equally convenient.
Evolution by natural selection is not an idle hypothesis. The genetic variation on which selection acts is well understood in principle all the way down to the molecular level.
Natural selection is a blind and undirected consequence of the interaction between variation and the environment. Natural selection exists only in the continuous present of the natural world: it has no memory of its previous actions, no plans for the future, or underlying purpose.
... if variations useful to any organic being do occur, assuredly individuals thus characterized will have the best chance of being perserved in the struggle for life; and from the strong principle of inheritance they will tend to produce offsping similarly characterized. This principle of preservation, I have called, for the sake of brevity, Natural Selection.
The essence of Darwinism lies in its claim that natural selection creates the fit. Variation is ubiquitous and random in direction. It supplies raw material only. Natural selection directs the course of evolutionary change.
Quantitative work shows clearly that natural selection is a reality, and that, among other things, it selects Mendelian genes, which are known to be distributed at random through wild populations, and to follow the laws of chance in their distribution to offspring. In other words, they are an agency producing variation of the kind which Darwin postulated as the raw material on which selection acts.
You put three facts together - that all organisms produce more offspring that can survive, that there's variation among organisms, and that at least some of that variation is inherited - and the syllogistic inference is natural selection.
The growth of our knowledge is the result of a process closely resembling what Darwin called 'natural selection'; that is, the natural selection of hypotheses: our knowledge consists, at every moment, of those hypotheses which have shown their (comparative) fitness by surviving so far in their struggle for existence, a competitive struggle which eliminates those hypotheses which are unfit.
Natural selection is just three factors - over-production, variation, and inheritance combined to produce adaptation to changing local environments. It's not a principle or progress; it's just a principle of local adaptation. You don't make better creatures in any cosmic sense; you make creatures that are better suited to the changing climates of their local habitats.
It may be said that natural selection is daily and hourly scrutinizing, throughout the world, every variation, even the slightest; rejecting that which is bad, preserving and adding up all that is good; silently and insensibly working, wherever and whenever opportunity offers, at the improvement of each organic being in relation to its organic and inorganic conditions of life.
Now let me step back from the problem and very generally discuss natural selection and what we know about it. I think it is safe to say that we know for sure that natural selection, as a process, does work. There is a mountain of experimental and observational evidence, much of it predating genetics, which shows that natural selection as a biological process works.
That there is much suffering in the world no one disputes. Which is more likely, that pain and evil are the result of an all-powerful and good God, or the product of uncaring natural forces? The presence of much suffering agrees well with the view that all organic beings have been developed through variation and natural selection.
This fundamental subject of Natural Selection will be treated at some length in the fourth chapter; and we shall then see how Natural Selection almost inevitably causes much Extinction of the less improved forms of life and induces what I have called Divergence of Character.
You can be a thorough-going Neo-Darwinian without imagination, metaphysics, poetry, conscience, or decency. For 'Natural Selection' has no moral significance: it deals with that part of evolution which has no purpose, no intelligence, and might more appropriately be called accidental selection, or better still, Unnatural Selection, since nothing is more unnatural than an accident. If it could be proved that the whole universe had been produced by such Selection, only fools and rascals could bear to live.
This preservation of favourable variations and the destruction of injurious variations, I call Natural Selection, or the Survival of the Fittest. Variations neither useful nor injurious would not be affected by natural selection and would be left a fluctuating element.
The basic formulation, or bare-bones mechanics, of natural selection is a disarmingly simple argument, based on three undeniable facts (overproduction of offspring, variation, and heritability) and one syllogistic inference (natural selection, or the claim that organisms enjoying differential reproductive success will, on average, be those variants that are fortuitously better adapted to changing local environments, and that these variants will then pass their favored traits to offspring by inheritance).
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